Antibodies & Immune Memory

01Antibodies & Immune Memory

Antibody structure

This section covers antibody structure, antigen-binding sites and the structural basis of antibody specificity.

Structure of an antibody

Antibody: a globular glycoprotein with antigen-binding sites that are specific to a particular antigen.

An antibody is built from four polypeptide chains: a pair of heavy chains and a pair of light chains. Disulfide bonds join these chains to produce the characteristic Y-shaped quaternary structure.

The molecule contains regions with different roles:

  • The variable regions form the antigen-binding sites near the ends of the two arms.
  • Differences in the amino acid sequence of the variable regions give different antibodies differently shaped antigen-binding sites.
  • The constant region is the same for antibodies within a particular class and is associated with the way the antibody brings about destruction.
  • Where present, a hinge region in the heavy chains gives flexibility so the binding sites can attach to antigens at different angles.

Each antibody has two antigen-binding sites, one at the end of each arm.

antigen-binding sitesvariableregionlight chainconstant regionheavy chainhinge regiondisul¯debonds2 heavy chains + 2 light chains

Specificity

The part of an antigen recognised by an antibody is called an epitope. An antigen-binding site can bind when its shape is complementary to that epitope.

Because the amino acid sequence of the variable region differs between antibodies, their antigen-binding sites also differ. This gives each antibody its particular antigen specificity.

Exam Tip: When explaining antibody specificity, link the sequence clearly: different amino acid sequence in the variable region → different antigen-binding-site shape → only a complementary antigen or epitope binds.

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